Augustynolophus is an extinct genus of herbivorous saurolophine hadrosaur dinosaur which was discovered in the Moreno Fo
Augustynolophus

Augustynolophus is an extinct genus of herbivorous saurolophine hadrosaur dinosaur which was discovered in the Moreno Formation in California, dating to the late Maastrichtian age, making it one of the last dinosaurs known from the fossil record before the Cretaceous–Paleogene extinction event.
Augustynolophus Temporal range: Late Cretaceous, | |
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Holotype skull, Natural History Museum of Los Angeles County | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | †Ornithischia |
Clade: | †Ornithopoda |
Family: | †Hadrosauridae |
Subfamily: | †Saurolophinae |
Genus: | †Augustynolophus Prieto-Márquez et al., 2014 |
Species: | †A. morrisi |
Binomial name | |
†Augustynolophus morrisi (Prieto-Márquez & Wagner, 2013) | |
Synonyms | |
Saurolophus morrisi Prieto-Márquez & Wagner, 2013 |
History of discovery

The generic name derives from a combination of the Augustyn family, who helped support the Los Angeles County Museum, and the suffix "-lophus," meaning crest, a reference to its relative Saurolophus. The specific name refers to palaeontologist William Morris. It was originally described as a species of Saurolophus, S. morrisi. However, when a more in-depth study took place, the end results revealed that its cranial structure was vastly different when it was juxtaposed with the other known members of the tribe Saurolophini, most notably Saurolophus osborni and Saurolophus angustirostris and Prosaurolophus maximus and therefore, it was determined to be a separate genus.
All known specimens of Augustynolophus have been found only in California, which was a part of Laramidia, among the best locations for dinosaur fossils. There are currently two known specimens of Augustynolophus. The holotype, LACM/CIT 2852, was unearthed in 1943. It consisted of the majority of the skull (including the dentary and predentary), vertebrae, and bones of the limb and hand. The second specimen was designated LACM/CIT 2760. Discovered in 1939, it was made up of elements of the skull and limbs. Due to its smaller size, it may have been a juvenile. It is one of three named dinosaurs from the western coast of the United States, the other two being the Campanian Aletopelta coombsi, and the sea bird Ichthyornis. Additionally there are a number of fragmentary remains which have been found of dinosaurs in all of the western coastal states, California, Oregon, and Washington. In addition to a tyrannosaur from Sucia Island of Washington, fragmentary remains of indeterminate hadrosaurs and ornithopods have been found throughout California as far back as the 1930s and more recently Oregon as well. In September 2017, Augustynolophus was declared the official state dinosaur of California.
Description

Augustynolophus was a large hadrosaur, reaching 8 metres (26 ft) in length and 3 metric tons (3.3 short tons) in body mass. Like all species of hadrosaur, Augustynolophus morrisi was a herbivorous dinosaur which had a diet consisting of the plant life in the area, it was specialized to chew its food since hadrosaurs were one of the few known species of dinosaur that chewed its food.
Paleoecology

Very is little is known about the fauna found in the coastal states of Laramida, which is a stark similarity to the neighboring island continent of Appalachia which was on the opposite side of the Western Interior Seaway. The western United States has a rich history of rich fossil finds, most notable examples include the Hell Creek Formation and the Two Medicine Formation. However, like with Appalachia, the land fauna of western Laramidia, most notably the dinosaurs, is not well studied and are not well known.
However, despite the fact that dinosaur fossils are rare in California, the Moreno Formation is one of the more well studied Mesozoic geological formations in California and like Appalachia, it is more well known for the large amount of marine fossils.Augustynolophus shared its environment with a wide variety of marine creatures which consisted of non-marine turtles, mosasaurs, plesiosaurs and ray-finned fish. The turtles that lived in this region included Basilemys and Osteopygis. Mosasaurs were very common in this region and 4 species have been unearthed from this fossil formation. They include Prognathodon, Halisaurus, Plesiotylosaurus, and Plotosaurus. Plesiosaurs that inhabited the Moreno Formation consisted of 4 genera: Frensosaurus, Morenosaurus, Aphrosaurus, and Hydrotherosaurus.Bonnerichthys and Saurodon were the only ray-finned fish found here. Indeterminate hadrosaur fossils are common in California too.
See also
- Timeline of hadrosaur research
- List of California state symbols
References
- Sues, H.D.; Bell, P.R.; Evans, D.C. (2010). "Revision of the status of Saurolophus (Hadrosauridae) from California, USA". Canadian Journal of Earth Sciences. 47 (11): 1417–1426. Bibcode:2010CaJES..47.1417S. doi:10.1139/e10-062.
- Lehman, T.M. (1987). "Late Maastrichtian paleoenvironments and dinosaur biogeography in the western interior of North America". Palaeogeography, Palaeoclimatology, Palaeoecology. 60: 189–217. Bibcode:1987PPP....60..189L. doi:10.1016/0031-0182(87)90032-0.
- Prieto-Márquez, Albert; Wagner, Jonathan R.; Bell, Phil R.; Chiappe, Luis M. (2014). "The late-surviving 'duck-billed' dinosaur Augustynolophus from the upper Maastrichtian of western North America and crest evolution in Saurolophini". Geological Magazine. 152 (2): 225–241. Bibcode:2015GeoM..152..225P. doi:10.1017/S0016756814000284. S2CID 131049979.
- Morris, W.J. (1982). "California dinosaurs". In Bottjer, DJ; Colburn, IP; Cooper, JD (eds.). Late Cretaceous Depositional Environments and Paleogeography, Santa Ana Mountains, Southern California. Los Angeles: Pacific Section, Society of Economic Paleontologists and Mineralogists.
- ; (2013). "A new species of saurolophine hadrosaurid dinosaur from the Late Cretaceous of the Pacific coast of North America". Acta Palaeontologica Polonica. 58 (2): 255–268. doi:10.4202/app.2011.0049.
- Brown, B. (1912). "A crested dinosaur from the Edmonton Cretaceous". Bulletin of the American Museum of Natural History: 131–136.
{{cite journal}}
: Cite journal requires|journal=
(help) - Brown, B. (1913). "The skeleton of Saurolophus, a crestedduck-billed dinosaur from the Edmonton Cretaceous". Bulletin of the American Museum of Natural History. 32: 387–393.
- Bell, P.R. (February 2011). "Redescription of the skull of Saurolophus osborni Brown 1912 (Ornithischia: Hadrosauridae)". Cretaceous Research. 32 (1): 30–44. Bibcode:2011CrRes..32...30B. doi:10.1016/j.cretres.2010.10.002.
- Bell, P.R. (3 February 2012). "Standardized terminology and potential taxonomic utility for hadrosaurid skin impressions: a case study for Saurolophus from Canada and Mongolia". PLOS ONE. 7 (2): e31295. Bibcode:2012PLoSO...731295B. doi:10.1371/journal.pone.0031295. PMC 3272031. PMID 22319623.
- Bell, P.R. (2011). "Cranial Osteology and Ontogeny of Saurolophus angustirostris from the late Cretaceous of Mongolia with comments on Saurolophus osborni from Canada". Acta Palaeontologica Polonica. 56 (4): 703–722. doi:10.4202/app.2010.0061.
- McGarrity, Christopher T.; Campione, Nicolas E.; Evans, David C. (27 March 2013). "Cranial anatomy and variation in Prosaurolophus maximus (Dinosauria: Hadrosauridae)". Zoological Journal of the Linnean Society. 167 (4): 531–568. doi:10.1111/zoj.12009. S2CID 83747340.
- Scott D. Sampson; Mark A. Loewen; Andrew A. Farke; Eric M. Roberts; Catherine A. Forster; Joshua A. Smith; Alan L. Titus (2010). Stepanova, Anna (ed.). "New Horned Dinosaurs from Utah Provide Evidence for Intracontinental Dinosaur Endemism". PLOS ONE. 5 (9): e12292. Bibcode:2010PLoSO...512292S. doi:10.1371/journal.pone.0012292. PMC 2929175. PMID 20877459.
- Hilton, Richard (2003). Dinosaurs and Other Mesozoic Reptiles of California. University of California Press. p. 264. ISBN 9780520233157.
- Ford, T.L. (1 October 2001). 12. Carlsbad Ankylosaur (Ornithischia, Ankylosauria): An Ankylosaurid and Not a Nodosaurid.= (1st ed.). Bloomington, Indiana: Indiana University Press. pp. 239–258. ISBN 978-0253339645. Retrieved 22 September 2016.
- Hilton RP, Gohre ES, Embree PG, Stidham TA. California’s first fossil evidence of Cretaceous winged vertebrates. California Geology 1999;52(4):4–10.
- Peecook, Brandon; Sidor, Christian (May 2015). "The First Dinosaur from Washington State and a Review of Pacific Coast Dinosaurs from North America". PLOS ONE. 10 (5): e0127792. Bibcode:2015PLoSO..1027792P. doi:10.1371/journal.pone.0127792. PMC 4439161. PMID 25993090.
- Rantz, Jason (January 13, 2021). "Renewed push to designate Suciasaurus rex the WA state dinosaur". KTTH AM 770.
- Morris, W.J. (May 1973). "A review of Pacific coast hadrosaurs". Journal of Paleontology. 47 (3): 551–561.
- HARPER, HILLIARD (20 May 1987). "Dinosaur Skeleton Found at Carlsbad Site". Los Angeles Times. Retrieved 24 September 2016.
- Ford, Tracy (January 1999). "A Review of the Hadrosaurs from California". Mesa Southwest Museum Bulletin. 6: 59–60.
- Stock, Chester (December 1939). "Occurrence of Cretaceous Reptiles in the Moreno Shales of the Southern Coast Ranges, California". Proceedings of the National Academy of Sciences. 25 (12): 617–620. Bibcode:1939PNAS...25..617S. doi:10.1073/pnas.25.12.617. ISSN 0027-8424. PMC 1077983. PMID 16588308.
- Black, Riley (2016-05-17). "The Dinosaurs of California's Lost World". KCET. Retrieved 2022-07-25.
- Hesse, Curtis J.; Welles, S. P. (1936-08-14). "The First Record of a Dinosaur from the West Coast". Science. 84 (2172): 157–158. doi:10.1126/science.84.2172.157-b. ISSN 0036-8075. S2CID 241227238.
- Retallack et al., 2018
- Strommer, Kristin (July 14, 2021). "UO-led team uncovers new pieces of Oregon's dinosaur puzzle". University of Oregon. Retrieved July 15, 2021.
- Taylor, David G.; Lucas, Spencer G. (2018). "A Late Cretaceous (Campanian) hadrosaur sacrum from the Cape Sebastian Sandstone, Curry County, Oregon". New Mexico Museum of Natural History and Science Bulletin. 79: 695–702.
- Blume, Howard (23 September 2017). "California gets its own official state dinosaur". Los Angeles Times.
- Paul, Greg (2010). The Princeton Field Guide to Dinosaurs. New Jersey: Princeton University Press. p. 335.
- Galton, P.M. (1973). "The cheeks of ornithischian dinosaurs". Lethaia. 6 (1): 67–89. Bibcode:1973Letha...6...67G. doi:10.1111/j.1502-3931.1973.tb00873.x.
- Tan, X.; Kodama, K.P. (1998). "Compaction-corrected inclinations from southern California Cretaceous marine sedimentary rocks indicate no paleolatitudinal offset for the Peninsular Ranges terrane". Journal of Geophysical Research: Solid Earth. 103 (B11): 27169–27192. Bibcode:1998JGR...10327169T. doi:10.1029/98jb02343.
- Switek, Brian (2016-05-17). "The Dinosaurs of California's Lost World". KCET. Retrieved 17 May 2016.
- Ford, K.W. (2006). "Stratigraphic positions of marine reptile and dinosaur specimens in the Moreno Formation, in the Tumey Hills and Panoche Hills, Fresno County, California". New Mexico Museum of Natural History and Science Bulletin. 35: 407–410.
- Popenoe, W.P. (1973). "Southern California Cretaceous formations and faunas with especial reference to the Simi Hills and Santa Monica Mountains". Cretaceous Stratigraphy of the Santa Monica Mountains and Simi Hills, Southern California. Pacific Section, SEPM, Geological Guidebook, Los Angeles, California: 15–20.
- Reeside, J.B. (1957). "Paleoecology of the Cretaceous seas of the Western Interior of the United States". Geological Society of America Memoirs. 67: 505–542. doi:10.1130/MEM67V2-p505.
- Stock, C. (1939). "Occurrence of Cretaceous reptiles in the Moreno shales of the Southern Coast Ranges, California" (PDF). Proceedings of the National Academy of Sciences. 25 (12): 617–620. Bibcode:1939PNAS...25..617S. doi:10.1073/pnas.25.12.617. PMC 1077983. PMID 16588308.
- Lyman, Theophan (2022). "Taphonomy of Marine Reptiles and Dinosaurs of the Upper Cretaceous Moreno Formation, California". Geological Society of America Abstracts with Programs. 54. Geological Society of America. Bibcode:2022GSAA...5476567L. doi:10.1130/abs/2022am-376567.
- Ford, Kerry (2006). "Stratigraphic positions of marine reptile and dinosaur specimens in the Moreno Formation, in the Tumey Hills and Panoche Hills, Fresno County, California". New Mexico Museum of Natural History and Science Bulletin. 35: 407–410.
- E., Foster, David (1980). Osteopygis Sp., a marine turtle from the late Cretaceous Moreno formation of California. Museum of Paleontology, University of California. OCLC 833332950.
{{cite book}}
: CS1 maint: multiple names: authors list (link) - LINDGREN, JOHAN; SCHULP, ANNE S. (September 2010). "New material ofPrognathodon(Squamata: Mosasauridae), and the mosasaur assemblage of the Maastrichtian of California, U.S.A.". Journal of Vertebrate Paleontology. 30 (5): 1632–1636. Bibcode:2010JVPal..30.1632L. doi:10.1080/02724634.2010.501444. ISSN 0272-4634. S2CID 131124198.
- O’Gorman, Jose P. (2019-09-03). "Elasmosaurid phylogeny and paleobiogeography, with a reappraisal of Aphrosaurus furlongi from the Maastrichtian of the Moreno Formation". Journal of Vertebrate Paleontology. 39 (5): e1692025. Bibcode:2019JVPal..39E2025O. doi:10.1080/02724634.2019.1692025. ISSN 0272-4634. S2CID 215756238.
- O'Gorman, José P. (December 2016). "Reappraisal of Fresnosaurus drescheri (Plesiosauria; Elasmosauridae) from the Maastrichtian Moreno Formation, California, USA". Cretaceous Research. 68: 9–20. Bibcode:2016CrRes..68....9O. doi:10.1016/j.cretres.2016.07.009. hdl:11336/54627. ISSN 0195-6671.
- Friedman, Matt; Shimada, Kenshu; Everhart, Michael J.; Irwin, Kelly J.; Grandstaff, Barbara S.; Stewart, J. D. (January 2013). "Geographic and stratigraphic distribution of the Late Cretaceous suspension-feeding bony fishBonnerichthys gladius(Teleostei, Pachycormiformes)". Journal of Vertebrate Paleontology. 33 (1): 35–47. Bibcode:2013JVPal..33...35F. doi:10.1080/02724634.2012.713059. ISSN 0272-4634. S2CID 128958842.
- Morris, William J. (1973). "A Review of Pacific Coast Hadrosaurs". Journal of Paleontology. 47 (3): 551–561. ISSN 0022-3360. JSTOR 1303203.
- Stock, Chester (1939). "Occurrence of Cretaceous Reptiles in the Moreno Shales of the Southern Coast Ranges, California". Proceedings of the National Academy of Sciences of the United States of America. 25 (12): 617–620. Bibcode:1939PNAS...25..617S. doi:10.1073/pnas.25.12.617. ISSN 0027-8424. JSTOR 87045. PMC 1077983. PMID 16588308.
- Harper, Hilliard (1987-05-20). "Dinosaur Skeleton Found at Carlsbad Site". Los Angeles Times. Retrieved 2025-03-06.
- Hesse, Curtis J.; Welles, S. P. (1936-08-14). "The First Record of a Dinosaur from the West Coast". Science. 84 (2172): 157–158. doi:10.1126/science.84.2172.157.b. ISSN 0036-8075.
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Augustynolophus is an extinct genus of herbivorous saurolophine hadrosaur dinosaur which was discovered in the Moreno Formation in California dating to the late Maastrichtian age making it one of the last dinosaurs known from the fossil record before the Cretaceous Paleogene extinction event Augustynolophus Temporal range Late Cretaceous 70 66 Ma PreꞒ Ꞓ O S D C P T J K Pg N Holotype skull Natural History Museum of Los Angeles CountyScientific classificationDomain EukaryotaKingdom AnimaliaPhylum ChordataClade DinosauriaClade OrnithischiaClade OrnithopodaFamily HadrosauridaeSubfamily SaurolophinaeGenus Augustynolophus Prieto Marquez et al 2014Species A morrisiBinomial name Augustynolophus morrisi Prieto Marquez amp Wagner 2013 SynonymsSaurolophus morrisi Prieto Marquez amp Wagner 2013History of discoveryPectoral and limb elements The generic name derives from a combination of the Augustyn family who helped support the Los Angeles County Museum and the suffix lophus meaning crest a reference to its relative Saurolophus The specific name refers to palaeontologist William Morris It was originally described as a species of Saurolophus S morrisi However when a more in depth study took place the end results revealed that its cranial structure was vastly different when it was juxtaposed with the other known members of the tribe Saurolophini most notably Saurolophus osborni and Saurolophus angustirostris and Prosaurolophus maximus and therefore it was determined to be a separate genus All known specimens of Augustynolophus have been found only in California which was a part of Laramidia among the best locations for dinosaur fossils There are currently two known specimens of Augustynolophus The holotype LACM CIT 2852 was unearthed in 1943 It consisted of the majority of the skull including the dentary and predentary vertebrae and bones of the limb and hand The second specimen was designated LACM CIT 2760 Discovered in 1939 it was made up of elements of the skull and limbs Due to its smaller size it may have been a juvenile It is one of three named dinosaurs from the western coast of the United States the other two being the Campanian Aletopelta coombsi and the sea bird Ichthyornis Additionally there are a number of fragmentary remains which have been found of dinosaurs in all of the western coastal states California Oregon and Washington In addition to a tyrannosaur from Sucia Island of Washington fragmentary remains of indeterminate hadrosaurs and ornithopods have been found throughout California as far back as the 1930s and more recently Oregon as well In September 2017 Augustynolophus was declared the official state dinosaur of California DescriptionLife restoration Augustynolophus was a large hadrosaur reaching 8 metres 26 ft in length and 3 metric tons 3 3 short tons in body mass Like all species of hadrosaur Augustynolophus morrisi was a herbivorous dinosaur which had a diet consisting of the plant life in the area it was specialized to chew its food since hadrosaurs were one of the few known species of dinosaur that chewed its food PaleoecologyFacial and mandibular elements Very is little is known about the fauna found in the coastal states of Laramida which is a stark similarity to the neighboring island continent of Appalachia which was on the opposite side of the Western Interior Seaway The western United States has a rich history of rich fossil finds most notable examples include the Hell Creek Formation and the Two Medicine Formation However like with Appalachia the land fauna of western Laramidia most notably the dinosaurs is not well studied and are not well known However despite the fact that dinosaur fossils are rare in California the Moreno Formation is one of the more well studied Mesozoic geological formations in California and like Appalachia it is more well known for the large amount of marine fossils Augustynolophus shared its environment with a wide variety of marine creatures which consisted of non marine turtles mosasaurs plesiosaurs and ray finned fish The turtles that lived in this region included Basilemys and Osteopygis Mosasaurs were very common in this region and 4 species have been unearthed from this fossil formation They include Prognathodon Halisaurus Plesiotylosaurus and Plotosaurus Plesiosaurs that inhabited the Moreno Formation consisted of 4 genera Frensosaurus Morenosaurus Aphrosaurus and Hydrotherosaurus Bonnerichthys and Saurodon were the only ray finned fish found here Indeterminate hadrosaur fossils are common in California too See alsoTimeline of hadrosaur research List of California state symbolsReferencesSues H D Bell P R Evans D C 2010 Revision of the status of Saurolophus Hadrosauridae from California USA Canadian Journal of Earth Sciences 47 11 1417 1426 Bibcode 2010CaJES 47 1417S doi 10 1139 e10 062 Lehman T M 1987 Late Maastrichtian paleoenvironments and dinosaur biogeography in the western interior of North America Palaeogeography Palaeoclimatology Palaeoecology 60 189 217 Bibcode 1987PPP 60 189L doi 10 1016 0031 0182 87 90032 0 Prieto Marquez Albert Wagner Jonathan R Bell Phil R Chiappe Luis M 2014 The late surviving duck billed dinosaur Augustynolophus from the upper Maastrichtian of western North America and crest evolution in Saurolophini Geological Magazine 152 2 225 241 Bibcode 2015GeoM 152 225P doi 10 1017 S0016756814000284 S2CID 131049979 Morris W J 1982 California dinosaurs In Bottjer DJ Colburn IP Cooper JD eds Late Cretaceous Depositional Environments and Paleogeography Santa Ana Mountains Southern California Los Angeles Pacific Section Society of Economic Paleontologists and Mineralogists 2013 A new species of saurolophine hadrosaurid dinosaur from the Late Cretaceous of the Pacific coast of North America Acta Palaeontologica Polonica 58 2 255 268 doi 10 4202 app 2011 0049 Brown B 1912 A crested dinosaur from the Edmonton Cretaceous Bulletin of the American Museum of Natural History 131 136 a href wiki Template Cite journal title Template Cite journal cite journal a Cite journal requires journal help Brown B 1913 The skeleton of Saurolophus a crestedduck billed dinosaur from the Edmonton Cretaceous Bulletin of the American Museum of Natural History 32 387 393 Bell P R February 2011 Redescription of the skull of Saurolophus osborni Brown 1912 Ornithischia Hadrosauridae Cretaceous Research 32 1 30 44 Bibcode 2011CrRes 32 30B doi 10 1016 j cretres 2010 10 002 Bell P R 3 February 2012 Standardized terminology and potential taxonomic utility for hadrosaurid skin impressions a case study for Saurolophus from Canada and Mongolia PLOS ONE 7 2 e31295 Bibcode 2012PLoSO 731295B doi 10 1371 journal pone 0031295 PMC 3272031 PMID 22319623 Bell P R 2011 Cranial Osteology and Ontogeny of Saurolophus angustirostris from the late Cretaceous of Mongolia with comments on Saurolophus osborni from Canada Acta Palaeontologica Polonica 56 4 703 722 doi 10 4202 app 2010 0061 McGarrity Christopher T Campione Nicolas E Evans David C 27 March 2013 Cranial anatomy and variation in Prosaurolophus maximus Dinosauria Hadrosauridae Zoological Journal of the Linnean Society 167 4 531 568 doi 10 1111 zoj 12009 S2CID 83747340 Scott D Sampson Mark A Loewen Andrew A Farke Eric M Roberts Catherine A Forster Joshua A Smith Alan L Titus 2010 Stepanova Anna ed New Horned Dinosaurs from Utah Provide Evidence for Intracontinental Dinosaur Endemism PLOS ONE 5 9 e12292 Bibcode 2010PLoSO 512292S doi 10 1371 journal pone 0012292 PMC 2929175 PMID 20877459 Hilton Richard 2003 Dinosaurs and Other Mesozoic Reptiles of California University of California Press p 264 ISBN 9780520233157 Ford T L 1 October 2001 12 Carlsbad Ankylosaur Ornithischia Ankylosauria An Ankylosaurid and Not a Nodosaurid 1st ed Bloomington Indiana Indiana University Press pp 239 258 ISBN 978 0253339645 Retrieved 22 September 2016 Hilton RP Gohre ES Embree PG Stidham TA California s first fossil evidence of Cretaceous winged vertebrates California Geology 1999 52 4 4 10 Peecook Brandon Sidor Christian May 2015 The First Dinosaur from Washington State and a Review of Pacific Coast Dinosaurs from North America PLOS ONE 10 5 e0127792 Bibcode 2015PLoSO 1027792P doi 10 1371 journal pone 0127792 PMC 4439161 PMID 25993090 Rantz Jason January 13 2021 Renewed push to designate Suciasaurus rex the WA state dinosaur KTTH AM 770 Morris W J May 1973 A review of Pacific coast hadrosaurs Journal of Paleontology 47 3 551 561 HARPER HILLIARD 20 May 1987 Dinosaur Skeleton Found at Carlsbad Site Los Angeles Times Retrieved 24 September 2016 Ford Tracy January 1999 A Review of the Hadrosaurs from California Mesa Southwest Museum Bulletin 6 59 60 Stock Chester December 1939 Occurrence of Cretaceous Reptiles in the Moreno Shales of the Southern Coast Ranges California Proceedings of the National Academy of Sciences 25 12 617 620 Bibcode 1939PNAS 25 617S doi 10 1073 pnas 25 12 617 ISSN 0027 8424 PMC 1077983 PMID 16588308 Black Riley 2016 05 17 The Dinosaurs of California s Lost World KCET Retrieved 2022 07 25 Hesse Curtis J Welles S P 1936 08 14 The First Record of a Dinosaur from the West Coast Science 84 2172 157 158 doi 10 1126 science 84 2172 157 b ISSN 0036 8075 S2CID 241227238 Retallack et al 2018 Strommer Kristin July 14 2021 UO led team uncovers new pieces of Oregon s dinosaur puzzle University of Oregon Retrieved July 15 2021 Taylor David G Lucas Spencer G 2018 A Late Cretaceous Campanian hadrosaur sacrum from the Cape Sebastian Sandstone Curry County Oregon New Mexico Museum of Natural History and Science Bulletin 79 695 702 Blume Howard 23 September 2017 California gets its own official state dinosaur Los Angeles Times Paul Greg 2010 The Princeton Field Guide to Dinosaurs New Jersey Princeton University Press p 335 Galton P M 1973 The cheeks of ornithischian dinosaurs Lethaia 6 1 67 89 Bibcode 1973Letha 6 67G doi 10 1111 j 1502 3931 1973 tb00873 x Tan X Kodama K P 1998 Compaction corrected inclinations from southern California Cretaceous marine sedimentary rocks indicate no paleolatitudinal offset for the Peninsular Ranges terrane Journal of Geophysical Research Solid Earth 103 B11 27169 27192 Bibcode 1998JGR 10327169T doi 10 1029 98jb02343 Switek Brian 2016 05 17 The Dinosaurs of California s Lost World KCET Retrieved 17 May 2016 Ford K W 2006 Stratigraphic positions of marine reptile and dinosaur specimens in the Moreno Formation in the Tumey Hills and Panoche Hills Fresno County California New Mexico Museum of Natural History and Science Bulletin 35 407 410 Popenoe W P 1973 Southern California Cretaceous formations and faunas with especial reference to the Simi Hills and Santa Monica Mountains Cretaceous Stratigraphy of the Santa Monica Mountains and Simi Hills Southern California Pacific Section SEPM Geological Guidebook Los Angeles California 15 20 Reeside J B 1957 Paleoecology of the Cretaceous seas of the Western Interior of the United States Geological Society of America Memoirs 67 505 542 doi 10 1130 MEM67V2 p505 Stock C 1939 Occurrence of Cretaceous reptiles in the Moreno shales of the Southern Coast Ranges California PDF Proceedings of the National Academy of Sciences 25 12 617 620 Bibcode 1939PNAS 25 617S doi 10 1073 pnas 25 12 617 PMC 1077983 PMID 16588308 Lyman Theophan 2022 Taphonomy of Marine Reptiles and Dinosaurs of the Upper Cretaceous Moreno Formation California Geological Society of America Abstracts with Programs 54 Geological Society of America Bibcode 2022GSAA 5476567L doi 10 1130 abs 2022am 376567 Ford Kerry 2006 Stratigraphic positions of marine reptile and dinosaur specimens in the Moreno Formation in the Tumey Hills and Panoche Hills Fresno County California New Mexico Museum of Natural History and Science Bulletin 35 407 410 E Foster David 1980 Osteopygis Sp a marine turtle from the late Cretaceous Moreno formation of California Museum of Paleontology University of California OCLC 833332950 a href wiki Template Cite book title Template Cite book cite book a CS1 maint multiple names authors list link LINDGREN JOHAN SCHULP ANNE S September 2010 New material ofPrognathodon Squamata Mosasauridae and the mosasaur assemblage of the Maastrichtian of California U S A Journal of Vertebrate Paleontology 30 5 1632 1636 Bibcode 2010JVPal 30 1632L doi 10 1080 02724634 2010 501444 ISSN 0272 4634 S2CID 131124198 O Gorman Jose P 2019 09 03 Elasmosaurid phylogeny and paleobiogeography with a reappraisal of Aphrosaurus furlongi from the Maastrichtian of the Moreno Formation Journal of Vertebrate Paleontology 39 5 e1692025 Bibcode 2019JVPal 39E2025O doi 10 1080 02724634 2019 1692025 ISSN 0272 4634 S2CID 215756238 O Gorman Jose P December 2016 Reappraisal of Fresnosaurus drescheri Plesiosauria Elasmosauridae from the Maastrichtian Moreno Formation California USA Cretaceous Research 68 9 20 Bibcode 2016CrRes 68 9O doi 10 1016 j cretres 2016 07 009 hdl 11336 54627 ISSN 0195 6671 Friedman Matt Shimada Kenshu Everhart Michael J Irwin Kelly J Grandstaff Barbara S Stewart J D January 2013 Geographic and stratigraphic distribution of the Late Cretaceous suspension feeding bony fishBonnerichthys gladius Teleostei Pachycormiformes Journal of Vertebrate Paleontology 33 1 35 47 Bibcode 2013JVPal 33 35F doi 10 1080 02724634 2012 713059 ISSN 0272 4634 S2CID 128958842 Morris William J 1973 A Review of Pacific Coast Hadrosaurs Journal of Paleontology 47 3 551 561 ISSN 0022 3360 JSTOR 1303203 Stock Chester 1939 Occurrence of Cretaceous Reptiles in the Moreno Shales of the Southern Coast Ranges California Proceedings of the National Academy of Sciences of the United States of America 25 12 617 620 Bibcode 1939PNAS 25 617S doi 10 1073 pnas 25 12 617 ISSN 0027 8424 JSTOR 87045 PMC 1077983 PMID 16588308 Harper Hilliard 1987 05 20 Dinosaur Skeleton Found at Carlsbad Site Los Angeles Times Retrieved 2025 03 06 Hesse Curtis J Welles S P 1936 08 14 The First Record of a Dinosaur from the West Coast Science 84 2172 157 158 doi 10 1126 science 84 2172 157 b ISSN 0036 8075